DeGilla was a clone of Admiral DeGill that he created to stand in for him until he could discover who was trying to assassinate him, however, DeGilla went rogue and teamed up with the assassin, who it turns out, was Pontifidora the Conquistadora (who he had fallen in love with upon first seeing her) to run the Galactic Guardians as their own private army. However, Pontifadora betrayed him and left him to die along with Betty and her crew, DeGilla helped them to escape and he took off, stating he would go to a place where he would be appreciated and start up his own army.
Due to a malfunction in the cloning machine, DeGilla is much stronger than DeGill and can morph into a much more bigger and muscular version of himself and speaks in a deeper voice in this form. DeGill had originally named the clone DeGill-A, but the clone changed it to DeGilla. DeGilla also has pointed teeth as opposed to DeGill's normal ones. ("Night of the DeGilla Monster")